Whole-Grain Spaghetti with Veggi-fied Meat Sauce

INGREDIENTS

6 oz

whole-grain spaghetti (such as Barilla)

Cooking spray

1 cup

finely chopped onion

8 oz

93% lean ground beef

garlic cloves, minced

1 cup

finely chopped zucchini

8 oz

cremini mushrooms, finely chopped

1 14.5 ounce can

unsalted diced tomatoes

1 tbsp

unsalted tomato paste

1/2 tsp

dried oregano

1/4 tsp

crushed red pepper

3/4 tsp

kosher salt

1/4 tsp

freshly ground black pepper

2 tbsp

finely grated Parmigiano-Reggiano cheese